The damage lies in a position that would allow a predator to sever attachment muscles and remove the body from the shell more easily. Lack of shell chips preserved adjacent to damaged ammonites precludes scavenging or post-burial crushing as causes of the damage. Ammonites were caught in the water column, manipulated into a horizontal position and then bitten at a precise point.
The most likely predator was an active swimmer, with the capability to hold and manipulate a smooth, possibly slippery, ammonite shell. Their conclusion was that the damage is consistent with bites made by robust, parrot-like, squid jaws.
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